Not sure how the air start was arranged on big Ruston singles, but their multi cylinder engines had a seperate sprung valve in each cylinder to admit air, 250 or 300 PSI, an air distributor to time the admission of air to each cylinder in sequence, and a sliding camshaft with 3 sets of cams, which allowed the intake valves to stay closed amd the exhaust valves to run on a double lobe cam to exhaust the cylinder every revolution, so it became a 2-stroke air motor for starting. There was also a half compression setting used during very cold starts while injecting fuel. Never known a failed start down to a few degrees below freezing, but it sure was not fun if the air tank had leaked down and you had to start the compressor engine first when near frozen. Combustor.